feather, the component structure of the outer covering and flight surfaces of all modern birds. Unique to birds, feathers apparently evolved from the scales of birds’ reptilian ancestors. The many different types of feathers are variously specialized for insulation, flight, formation of body contours, display, and sensory reception. Unlike the hair of most mammals, feathers do not cover the entire skin surface of birds but are arranged in symmetrical tracts (pterylae) with areas of bare skin (apteria) between. The latter may contain the small soft feathers called down. The typical feather consists of a central shaft (rachis), with serial paired